I have been revisiting the wetlands to hopefully watch the Virginia rail. I have been lucky enough to see it again! It has been venturing a little farther into the open, and I got to see it walking around and foraging. One of the things I noticed is that sometimes the rail used its whole foot as it walked through the wetlands.
Most birds are digitigrade animals which means that they walk around on their toes, not the entire foot. The backward-bending joint that looks like a knee is actually the bird's ankle. The rail mostly walked around on its toes. However, it often moved or stood in a kind of crouched position and engaged its whole foot. This seems like it would provide support with balance and stability while walking on slippery surfaces. I could also see it useful when stalking, lunging at prey, or leaning forward to probe around with its bill. Additionally, the rail has long toes which helps distribute its weight when walking on soft surfaces like the muddy bottom of a marsh. I imagine that long toes with claws further aid in stabilizing and gripping slippery stems and leaves of lilies like in the wetlands.